The species to
which you and all other living human beings on this planet belong
is Homo sapiens. Anatomically, modern humans can generally
be characterized by the lighter build of their skeletons compared
to earlier humans. Modern humans also have very large brains, which
vary in size from population to population and between males and
females, but the average is around 1300 cc. Housing this enlarged
brain has involved the reorganization of the skull into what is
thought of as the "modern" appearance -- a high vaulted
cranium with a flat and near vertical forehead. The supraorbital
torus is lost in most modern humans, and ridging above the orbits
in general is very reduced. The widest part of the skull is high
on the skull, as opposed to earlier Homo
erectus and H. ergaster. The
back of the skull lacks the transverse torus of H. erectus
and the occipital bun of H. neanderthalensis (Compare
the crania of H. neanderthalensis and H. sapiens).
The origin of
modern Homo sapiens is not yet resolved. Two extreme scenarios
have been proposed. According to the first, the distribution of
anatomical traits in modern human populations in different regions
was inherited from local populations of Homo erectus and
intermediate "archaic" forms. This "Multiregional
Hypothesis" states that all modern humans evolved in parallel
from earlier populations in Africa, Europe and Asia, with some genetic
intermixing among these regions. Support for this comes from the
similarity of certain minor anatomical structures in modern human
populations and preceding populations of Homo erectus in
the same regions.
A different model
proposes that a small, relatively isolated population of early humans
evolved into modern Homo sapiens, and that this population
succeeded in spreading across Africa, Europe, and Asia -- displacing
and eventually replacing all other early human populations as they
spread. In this scenario the variation among modern populations
is a recent phenomenon. Part of the evidence to support this theory
comes from molecular biology, especially studies of the diversity
and mutation rate of nuclear DNA and mitochondrial DNA in living
human cells.From these
studies an approximate time of divergence from the common ancestor
of all modern human populations can be calculated. This research
has typically yielded dates around 200,000 years ago, too young
for the "Multiregional Hypothesis." Molecular methods
have also tended to point to an African origin for all modern humans,
implying that the ancestral population of all living people migrated
from Africa to other parts of the world -- thus the name of this
interpretation: the "Out of Africa Hypothesis."
Whichever model
(if either) is correct, the oldest fossil evidence for anatomically
modern humans is about 130,000 years old in Africa, and there is
evidence for modern humans in the Near East sometime before 90,000
years ago.