Brian G. Richmond

Research Associate

Portrait of Brian Richmond.

Brian Richmond is Associate Professor in the Departments of Anthropology, and Anatomy and Regenerative Biology at the George Washington University, and Research Associate of the Human Origins Program, National Museum of Natural History, Smithsonian Institution.  He earned in B.S. in 1990 at Rice University and his Ph.D. in 1998 at Stony Brook University.  He is a Biological Anthropologist who works in the field to recover new fossils and in the lab to improve our abilities to interpret the fossil record, both of which are critical to our understanding of human evolution.  His research focuses on functional anatomy and human evolution, especially in the context of questions about the origin and evolution of human gait and the functional anatomy of the hand. His current projects include paleontological fieldwork in Koobi Fora, Kenya, the recovery and analysis of fossil footprints, developing new approaches to interpret behavior from skeletal anatomy, and investigating the anatomy and biomechanics of our earliest human ancestors and relatives.


Griffin N.L., Richmond B.G. 2010. Joint orientation and function in great ape and human proximal pedal phalanges. American Journal of Physical Anthropology 141, 116-123.

Bennett M.R., Harris J.W., Richmond B.G., Braun D.R., Mbua E., Kiura P., Olago D., Kibunjia M., Omuombo C., Behrensmeyer A.K., Huddart D., Gonzalez S. 2009. Early hominin foot morphology based on 1.5-million-year-old footprints from Ileret, Kenya. Science 323, 1197-1201.

Strait D.S., Weber G.W., Neubauer S., Chalk J., Richmond B.G., Lucas P.W., Spencer M.A., Schrein C., Dechow P.C., Ross C.F., Grosse I.R., Wright B.W., Constantino P., Wood B.A., Lawn B., Hylander W.L., Wang Q., Byron C., Slice D.E., Smith A.L. 2009. The feeding biomechanics and dietary ecology of Australopithecus africanus. Proceedings of the National Academy of Science U S A 106, 2124-2129.

Gordon A.D., Green D.J., Richmond B.G. 2008. Strong postcranial size dimorphism in Australopithecus afarensis: results from two new resampling methods for multivariate data sets with missing data. American Journal of Physical Anthropology 135, 311-328.

Griffin N.L., Gordon A.D., Richmond B.G., Anton S.C. 2008. Cross-sectional geometric analysis of a foot bone assemblage from Mangaia, Cook Islands. Homo 59, 27-40.

Lemelin P., Hamrick M.W., Richmond B.G., Godfrey L.R., Jungers W.L., Burney D.A. 2008. New hand bones of Hadropithecus stenognathus: implications for the paleobiology of the Archaeolemuridae. Journal of Human Evolution 54, 405-413.

Richmond B.G., Jungers W.L. 2008. Orrorin tugenensis femoral morphology and the evolution of hominin bipedalism. Science 319, 1662-1665.

Strait D.S., Wright B.W., Richmond B.G., Ross C.F., Dechow P.C., Spencer M.A., Wang Q. 2008 Craniofacial strain patterns during premolar loading: Implications for human evolution. In: Vinyard C. (ed) Primate Craniofacial Function and Biology. Springer, New York, pp 173-198.

Wang Q., Dechow P.C., Wright B.W., Ross C.F., Strait D.S., Richmond B.G., Spencer M.A., Byron C.D. 2008 Surface strain on bone and sutures in a monkey facial skeleton: an in vitro approach and its relevance to Finite Element Analysis.  In: Vinyard C. (ed) Primate Craniofacial Function and Biology. Springer, New York, pp 149-172.

Begun D.R., Richmond B.G., Strait D.S. 2007. Comment on "Origin of human bipedalism as an adaptation for locomotion on flexible branches". Science 318, 1066; author reply 1066.

Green D.J., Gordon A.D., Richmond B.G. 2007. Limb-size proportions in Australopithecus afarensis and Australopithecus africanus. Journal of Human Evolution 52, 187-200.

Richmond B.G. 2007. Biomechanics of phalangeal curvature. Journal of Human Evolution 53, 678-690.

Strait D.S., Richmond B.G., Spencer M.A., Ross C.F., Dechow P.C., Wood B.A. 2007. Masticatory biomechanics and its relevance to early hominid phylogeny: An examination of palatal thickness using finite-element analysis. Journal of Human Evolution 52, 585-599.

Richmond B.G. 2006 Functional morphology of the midcarpal joint in knuckle-walkers and terrestrial quadrupeds. In: Ishida H., Tuttle R., Pickford M., Ogihara N., Nakatsukasa M. (eds) Human Origins and Environmental Backgrounds. Springer, New York, pp 105-122.

Griffin N.L., Richmond B.G. 2005. Cross-sectional geometry of the human forefoot. Bone 37, 253-260.