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Ray Bernor

photo of Ray Bernor

Ray Bernor is Professor of Anatomy at Howard University, Laboratory of Evolutionary Biology.  He received his PhD at UCLA in 1978 with a major in Anthropology and minor in Vertebrate Paleontology.  Much of Ray’s work has been in middle and late Miocene faunas and biogeography of Eurasia and since 2000 has been especially engaged in late Miocene – Pleistocene human origins projects in Libya, Ethiopia, Kenya, Tanzania and South Africa.  Ray has a specialty in ungulate evolution and paleodiet, especially the evolutionary biology of fossil and living equids.

Selected Publications

Campbell, B.G., M.H. Amini, R.L. Bernor, W. Dickinson, R. Drake, R. Morris, J.A. Van Couvering and J.A.H. Van Couvering. 1980.  Maragheh: A Classical Late Miocene Vertebrate Locality in Northwestern Iran. Nature 287: 837-841.

Bernor, R.L. Geochronology and Zoogeographic Relationships of Miocene Hominoidea. 1983.   In: R.L. Ciochon and R. Corruccini, eds., New Interpretations of Ape and Human Ancestry.  Plenum Press, New York: 21-64.

Bernor, R.L., H. Tobien, L.-A. Hayek and H.-W. Mittmann.  1997.  The Höwenegg Hipparionine Horses: Systematics, Stratigraphy, Taphonomy and Paleoenvironmental Context.  Andrias 10: 1-230.

White, T., G. Suwa, B. Asfaw, S. Ambrose, Y. Beyene,  R. Bernor, J-R. Boisserie, B. Currie, H. Gilbert,  Y. Haile-Selassie, W. Hart, L. Hlusko, F.C. Howell, T. Reiko , R.T. Kono, A. Louchart, O. Lovejoy, P.  Renne, S. Haruo, E. Vrba, E., H.Wesselman, H., and G. Wolde-Gabriel .  2006.  Asa Issie, Aramis, and the Origin of Australopithecus.  Nature 440: 883-889.

Bernor, R.L.  New apes fill the gap. 2007.  Proceedings National Academy of Sciences USA. 104 (50): 19661-19662.

Eronen, J.T., M. Mirzaie, A. Karme. A. Micheels, R.L. Bernor and M. Fortelius.  Distribution History and Climatic Controls of the Late Miocene Pikermian Chronofauna.  2009.  Proceedings of the National Academy of Sciences. 106 (29): 11867-11871.  

White, T.D., S.H. Ambrose, G. Suwa, D. F. Su, D. DeGusta, R.L. Bernor, J.-R. Boisserie, M. Brunet, E. Delson, S. Frost, N. Garcia, I.X. Giaourtsakis, Y. Haile-Selassie, F. C. Howell, T. Lehmann, A. Likius, C. Pehlevan, H. Saegusa, G. Semprebon, M. Teaford and E. Vrba.  2009.  Macrovertebrate Paleontology and the Pliocene Habitat of Ardipithecus ramidus.  Science  326:87-93.

Bernor, R.L., M. Armour-Chelu, H. Gilbert, T. Kaiser and E. Schulz.  2010.  Equidae., pp. 685-721; L. Werdelin and B. Sanders (eds.) Cenozoic Mammals of Africa. University of California Press. 

Wolf, D., Bernor, R.L.* and S.T. Hussain*.  2013. Evolution, Biostratigraphy and Geochronology of Siwalik Hipparionine Horses. Palaeontographica.  300: 1-118.

Suwa, G., Y. Beyene, H. Nakaya, R.L. Bernor, J.-R. Boisserie, F. Bibi, S.H. Ambrose, K. Sano, S. Katoh and B. Asfaw.  2015.  Newly discovered cercopithecid, equid and other mammalian fossils from the Chorora Formation, Ethiopia. Anthropology Science 123(1): 19-39.

Katoh, S., Y. Beyene, T. Itaya, H. Hyodo, M. Hyodo, K. Yagi, C. Gouzu, G. WoldeGabriel, W.K. Hart, S.H. Ambrose, H. Nakaya, R.L. Bernor, J.-R. Boisseri, F. Bibi, Haruo. Saegusa, T. Sasaki, K. Sano, B. Asfaw and G. Suwa.  2016.  New Geological and paleontological age constraint for the gorilla-human lineage split.  Nature 530 (No. 7580; February 11): 215-218.  doi:10.1038/nature16510.

Edited Volumes

Bernor, R.L., V. Fahlbusch and H.-W. Mittmann (eds.) 1996. The Evolution of Western Eurasian Neogene Mammal Faunas; Columbia University Press, New York. 487 pp.

Fortelius, M. J. Kappelman, S. Sen and R.L. Bernor (eds.) 2003.  The Geology and Paleontology of the Miocene Sinap Formation, Turkey. Columbia University Press: New York.  409 pp.

Bernor, R.L., L. Kordos and L. Rook.  2005.  Multidisciplinary Research at Rudabánya; in: R.L. Bernor, L. Kordos and L. Rook (eds.): Multidisciplinary Research at Rudabánya. Paleontographica Italiana.  90: 1-313.